March of the Giant Penguins
Fossils Reveal Early Penguins Reaching 5 Ft. Tall Lived Near the Equator During One of Earth’s Warmest Periods
June 25, 2007
Media Contact: Tracey Peake, News Services, 919.515.6142 or tracey_peake@ncsu.edu
Reconstructions of the first Paleogene penguins from equatorial regions, illustrating morphological diversity and size range in present the new early penguin faunas. The late Eocene giant penguin Icadyptes salasi (right) and the middle Eocene Perudyptes devriesi (left) are shown to scale with the only extant penguin inhabiting Peru, Spheniscus humbolti (center). Icadyptes salasi is the first giant penguin known from a complete skull and had an estimated standing height of 1.5m. Perudyptes devriesi is known from one of the most complete skeletons of a more basal part of the penguin lineage. Together these new finds are revolutionizing our understanding of biogeography and timing of diversification in the penguin lineage. |
Giant prehistoric penguins? In Peru? It sounds more like something out of Hollywood than science, but a researcher from North Carolina State University along with U.S., Peruvian and Argentine collaborators has shown that two heretofore undiscovered penguin species reached equatorial regions tens of millions of years earlier than expected and during a period when the earth was much warmer than it is now.
Paleontologist Dr. Julia Clarke, assistant professor of marine, earth and atmospheric sciences at NC State with appointments at the North Carolina Museum of Natural Sciences and the American Museum of Natural History, and colleagues studied two newly discovered extinct species of penguins. Peruvian paleontologists discovered the new penguins’ sites in 2005.
The research is published online the week of June 25 in Proceedings of the National Academy of Sciences. It was funded by the National Science Foundation Office of International Science and Engineering and the National Geographic Society.
The first of the new species, Icadyptes salasi, stood 5 feet tall and lived about 36 million years ago. The second new species, Perudyptes devriesi, lived about 42 million years ago, was approximately the same size as a living King Penguin (2 ½ to 3 feet tall) and represents a very early part of penguin evolutionary history. Both of these species lived on the southern coast of Peru.
These new penguin fossils are among the most complete yet recovered and call into question hypotheses about the timing and pattern of penguin evolution and expansion. Previous theories held that penguins probably evolved in high latitudes (Antarctica and New Zealand) and then moved into lower latitudes that are closer to the equator about 10 million years ago – long after significant global cooling that occurred about 34 million years ago.
“We tend to think of penguins as being cold-adapted species,” Clarke says, “even the small penguins in equatorial regions today, but the new fossils date back to one of the warmest periods in the last 65 million years of Earth’s history. The evidence indicates that penguins reached low latitude regions more than 30 million years prior to our previous estimates.”
The new species are the first fossils to indicate a significant and diverse presence of penguins in equatorial areas during a period that predates one of the most important climatic shifts in Earth’s history, the transition from extremely warm temperatures in the Paleocene and Eocene Epochs to the development of “icehouse” Earth conditions and permanent polar icecaps. Not only did penguins reach low latitudes during this warmer interval, but they thrived: more species are known from the new Peruvian localities than inhabit those regions today.
By comparing the pattern of evolutionary relationships with the geographic distribution of other fossil penguins, Clarke and colleagues estimate that the two Peruvian species are the product of two separate dispersal events. The ancestors of Perudyptes appear to have inhabited Antarctica, while those of Icadyptes may have originated near New Zealand.
The first complete skull of a giant penguin. The skull is from the new species Icadyptes salasi (1.5m estimated standing height) from the late Eocene of Peru. Fine vascular texturing and other features of hyperelongate beak are unlike any extant or extinct penguin known. A skull of Spheniscus humbolti, the only species inhabiting Peru today, is shown for scale. Icadyptes salasi and a second new species described in this issue are revolutionizing our understanding of biogeography and diversity in the early penguin lineage. (Scale bar = 1cm). |
The new penguin specimens are among the most complete yet discovered that show us what early penguins looked like. Both new species had long narrow pointed beaks – now believed to be an ancestral beak shape for all penguins. Perudyptes devriesi has a slightly longer beak than seen in some living penguins but the giant Icadyptes salasi exhibits a grossly elongated beak with features not known in any extinct or living species. This species’ beak is sharply pointed, almost spear-like in appearance, and its neck is robustly built with strong muscle attachment sites. Icadyptes salasi is among the largest species of penguin yet described.
Although these fossils seem to contradict some of what we think we know about the relationship between penguins and climate, Clarke cautions against assuming that just because prehistoric penguins may not have been cold-adapted, living penguins won’t be negatively affected by climate change.
“These Peruvian species are early branches off the penguin family tree, that are comparatively distant cousins of living penguins,” Clarke says. “In addition, current global warming is occurring on a significantly shorter timescale. The data from these new fossil species cannot be used to argue that warming wouldn’t negatively impact living penguins.”
Note to Editors: An abstract of the PNAS paper follows
“Paleogene equatorial penguins challenge the proposed relationship between penguin biogeography, diversity, and Cenozoic climate change”
Authors: Julia A. Clarke, North Carolina State University; Daniel T. Ksepka, American Museum of Natural History; et al.
Published: Online the week of June 25, 2007, in Proceedings of the National Academy of Sciences
Abstract: New penguin fossils from the Eocene of Peru force a reevaluation of previous hypotheses regarding the causal role of climate change in penguin evolution. Repeatedly, it was proposed that penguins originated in high southern latitudes and arrived at equatorial regions relatively recently (e.g., 4 to 8 million years ago), only well after the onset of latest Eocene/Oligocene global cooling and increases in polar ice volume. By contrast, new discoveries from the middle and late Eocene of Peru reveal that penguins invaded low latitudes over 30 million years earlier than prior data supported, during one of the warmest intervals of the Cenozoic. A diverse fauna includes two new species, here reported from two of the best exemplars of Paleogene penguins yet recovered. The most comprehensive phylogenetic analysis of Sphenisciformes to date, combining morphological and molecular data, places the new species outside the extant penguin radiation (crown clade: Spheniscidae) and supports two separate dispersals to equatorial (paleolatitude ~14° S) regions during greenhouse Earth conditions. One new species is among the deepest divergences within Sphenisciformes. The second is the most complete giant (>1.5m standing height) penguin yet described. Both species provide critical information on early penguin cranial osteology, trends in penguin body size, and the evolution of the penguin flipper.
Dr. Julia Clarke is a research curator of paleontology for the Museum. She can be reached for interview at julia_clarke@ncsu.edu.
The North Carolina Museum of Natural Sciences, in downtown Raleigh, documents and interprets the natural history of the state of North Carolina through exhibits, research, collections, publications, and educational programming. Hours: Mon.-Sat., 9 am to 5 pm, and Sun., noon to 5 pm. Admission is free. Visit the Museum on the Web at naturalsciences.org. The Museum is an agency of the N.C. Department of Environment and Natural Resources, William G. Ross Jr., Secretary.
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